LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Ecuador released 158 juveniles in the first phase of an ambitious rewilding effort at the World Heritage Site.

PHILADELPHIA, Pa. — A pair of critically endangered, nearly 100-year-old Galapagos tortoises at the Philadelphia Zoo have become first-time parents. Subscribe to read this story ad-free Get unlimited ...