After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

Gramma, the beloved Galapagos tortoise who lived at the San Diego Zoo for nearly a century, has died on Nov. 20. She was (about) 141. The zoo's care team "made the compassionate and exceptionally ...

Gramma, a 141-year-old Galapagos tortoise and the San Diego Zoo’s oldest resident, has died. She came from the Bronx Zoo nearly a century ago and was known as the "Queen of the Zoo." Visitors and ...

LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.